#NEXUS [MacClade 4.08 registered to Susanne Renner, LMU Munich LMU Munich] BEGIN DATA; DIMENSIONS NTAX=111 NCHAR=81; FORMAT SYMBOLS= " 0 1 2 3 4 5 6 7 8 9" MISSING=N GAP=- ; CHARSTATELABELS 1 1._Flower_sexuality / bisexual unisexual, 2 2._Perigon / present absent, 3 3._Trichosclerids / absent 'present,_large,_not_in_bundles' 'present,_small,_in_bundles', 4 4._Spathe_structure / 'bract-like,_unspecialized,_�_inoconspicious' modified_for_attraction_or_specialized_in_some_other_way absent, 5 5._Inflorescence_stipe_and_peduncle / major_internode_of_infl._is_stipe_between_apadix_and_spathe major_internode_of_infl._is_peduncle_between_spathe_and_next_leaf_below_spathe, 6 6._Shoot_architecture / n_continuation_shoot_in_last_leaf_axil_before_spathe 'n-1_constinuation_shoot_in_axil_of_penultimate_leaf_before_spathe' culm_type_inflor.with_caulin_bracts_apart_from_spathe monopodial_scandent_structure '"no_shoot"', 7 7._Phyllotaxy / distichous spiral, 8 8._Pollen_aperture / monosulcate extended_monosulcate zonate diaperturate forate inaperturate monosulcoidate ulcerate multiaperturate, 9 9._Pollen_units / monads tetrads dyads, 10 10._Pollen_shape / ellipsoid 'hamburger-shaped' globose, 11 11._Mean_pollen_size / 'small_10-24�m' 'medium_25-49�m' 'large_50-99�m' 'very_large_100-199�m', 12 12._Pollen_exine_surface / reticulate_or_other smooth spinose striate, 13 13._Petiole_geniculum / absent present, 14 14._Pollen_starch / absent present, 15 15._Sclerotic_hypodermis_in_roots / absent present, 16 16._Endothecial_thickenings / present absent, 17 17._Resin_canals_in_roots / absent present, 18 18._Laticifers / absent_or_only_scattered_cells 'present,_simple,_articulated' 'present,_anastomosing', 19 19._Latex_s_bodys / absent present, 20 '20. Latex rubber (r) particles' / absent present, 21 21._Latex_y_bodys / absent present, 22 22._Latex_r_bodys / absent present, 23 23._Primary_leaf_venation / midrip_of_primary_vein_�_absent_with_veins_arcuate_from_the_base 'midrib_of_primary_veins_well_developed,_i.e._pinnate_veins_in_anterior_division', 24 '24. Primary leaf venation - basal ribs' / absent very_well_developed distinct_but_short, 25 '25. Primary leaf venation - blade' / 'distinct_blade_not_differentiated,_leaf_�_linear' distinct_blade_differentiated__ 'distinct_blade_not_differentiated,_leaf_anf_shoot_fused_into_thallus-like_body_called_frond', 26 '26. Primary leaf venation - marginal venation of anterior division or ultimate leaf lobes' / '�_no_sympodial_marginal_or_inframarginal_vein,_primary_veins_usually_fusing_near_apex' 'sympodial_marginal_vein_formed_of_�_all_primary_veins,_no_sympodial_inframarginal_vein' 'lowermost_primary_veins_forming_non-sympodial_marginal_veins,_sympodial_inframarginal_vein_formed_by_majority_of_primary_veins', 27 27._Marginal_form_of_leaf / not_lobed_individually_around_primary_veins lobed_pinnately_in_the_anterior_division lobed_pedately_or_radiately 'lobed_both_pinnately_and_pedately_(dracontioid)' lobed_trifidly_including_deeply_sagittate 'vi-_to_tri-_to_quadripinnate', 28 28._Fine_venation / secondaries_and_tertiaries_forming_mostly_cross_veins_to_primaries 'secondaries_and_tertiaries_parallel_to_primaries,_joined_by_cross_veins_only' 'secondaries_and_tertiaries_�_parallel_to_primaries,_often_forming_interprimary_sympodial_veins,_cross_connections_often_reticulate' 'secondaries_and_tertiaries_mostly_reticulating_freely,_forming_interprimary_sympodial_veins' 'absent,_only_primary_veins_present', 29 29._leaf_fenestration / fenestration_by_necrosis_between_primary_veins_absent fenestration_by_necrosis_between_primary_veins_present, 30 30._Prickles_on_stem_or_petiole / absent present, 31 31._Stem_producing_erect_shoots_with_bulbils / absent present, 32 '32. Stem type - thickening' / not_condensed_and_strongly_thickened 'condensed_,_strongly_thickened_but_not_depressend-globose' 'condensed,_strongly_thickened_into_depressed_globose_tuber_(corm)' 'stem_reduced_into_minute_button,_or_indistinguishable_from_thalloid_structure', 33 '33. Stem type - habit' / 'subareal,_creeping_to_erect_' '�_erect,_at_least_distally,_aerial' entirely_subterranean 'aerial_and_climbing_(incl._hemiepiphytes)' aerial_or_truly_epiphytic, 34 34._Peltate_leaves / absent present, 35 35._Aquatic_habit / not_aquatic helophytes floating_aquatics submerged_aquatics rheophytes, 36 36._Petiole_ligule / 'petile_sheath_not_lon-ligulate_apically' 'petiole_sheath_long-ligulate_apically', 37 37._Inframarginal_squamules / absent present, 38 38._Spathe_behaviour / 'no_differentiation_in_persistence,_entire_spathe_persistent_or_withering_slowly,_without_distinct_abscission' 'tube_or_lower_half_persistent,_blade_marescent_or_caducous' 'no_differentiation,_entire_spathe_soon_deciduous_or_marescent_with_distinct_basal_abscission', 39 39._Spathe_shape / 'fully_expanded,_often_reflexed' 'boat-shaped_�_convolute_basally_or_not' 'convolute_basally_into_distinct_tube_with_distinctly_different,_�_expanded_blade', 40 '40. Spadix-spathe fusion: chambers' / not_forming_2_distinct_chambers_by_fusion forming_2_distinct_chambers_by_partial_fusion, 41 '41. Spadix-spathe fusion: internal flap' / spathe_without_internal_flap_covering spathe_with_internal_flap_covering_and_adnate_to_spadix_apex, 42 42._Spathe_margins / free_or_connate_only_at_extreme_base connate_for_distinct_distance, 43 '43. Sapdix-fusion: dorsal fusion' / spadiy_dorsally_free_of_spathe spadix_female_zone_adnatea_to_spathe spadix_entirely_adnate_to_spathe, 44 44._Spadix_zonation / no 'female-male' 'female-sterile-male' 'female-male-sterile' 'female-sterile-male-sterile', 45 45._Spadix_appendix / absent_or_inconspicious 'conspicious_and_well_developed,_staminodial' 'conspicious_and_well_developed,_smooth_to_corrugate', 46 46._Basipetal_flowering / not_basipetal basipetal, 47 47._Female_zone_length / composed_of_more_than_1_flower composed_of_only_1_flower, 48 48._Stamen_thecae_horns / thecae_not_horned thecae_horned, 49 49._Stamen_connective / not_strongly_thickened 'strongly_thickened_laterally,_at_apex,__�_glandular', 50 50._Stamen_connation / free connate_by_filaments entiraly_connate, 51 51._Anther_dehiscence / by_longitudinal_slits 'by_oblique_pore-like_slits' by_apical_pores, 52 52._Staminodes_in_female_zone / absent present, 53 53._Ovary_locules / '2_-_3' unilocular '>_3', 54 54._Ovule_type / anatropous_or_hemianatropous campylotropous hemiorthotropous strictly_orthotropous, 55 55._Endosperm / 'copious,_embryo_relatively_small' 'sparse_to_absent,_embryo_relatively_large', 56 56._Placentation / axile_or_strongly_intrusive_placentae apical 'parietal,_septa_very_reduced' 'basal,_basal-parietal_or_basal-axile' baasal_and_apical, 57 57._Base_Chromosome_Number / 12 13 14 11 19 10 15 16 17 18, 58 58._Staminodes / 'absent,_prismatic_or_fungiform' 'hair-like,_subulate,_bristle-like,_clavate-elongated', 59 59._Style / not_laterally_thickened_or_extended 'laterally_thickened_or_extenden_intro_"mantel"_and_contiguous', 60 60._Ovule_number / '>_3_per_locule' '1_-_2_per_locule', 61 '61. Perigon-tepal connation' / tepals_free_or_partly_free perigon_a_single_unit, 62 '62. Perigon-tepal apex' / unthickened thickened, 63 63._Pistillodes / absent present, 64 64._Leaf_TS_spongy_aerenchym / 'type_1,_1a' type_2 type_3 type_4, 65 65._Leaf_TS_collenchyma / type_B type_Bi type_Sb type_Svc type_Sv absent, 66 66._Leaf_TS_vascular_bundles / 'broad,_phloem_strands_semi-circular,_type_1' 'narrow,_phloem_strands_elliptical,_type_2' 'small,_xylem_a_wide_cell_or_lacuna,_type_3', 67 67._Petiole_TS_collenchyma / type_B type_Bi type_Sb type_Svc type_Sv endheathing_vascular_bundles absent, 68 68._Petiole_TS_ground_tissue / cells_compact_in_centre 'type_1,_2' type_3 type_4, 69 69._Petiole_TS_vascular_bundles / 'large,_typ_1' 'medium-sized,_type_2' 'small,_with_dominant_single_xylem_element,_type_3', 70 70._Cotyledon_type / haustorial storage, 71 71._Cotyledon_shape / compact 'hypophyll_widened,_blade-like', 72 72._First_leaf / cataphyll eophyll, 73 73._Hypocotyl / present absent, 74 74._Primary_root / present absent, 75 75._Sporopollenin / present absent, 76 76._Roots / present absent, 77 77._Root_branching / present absent, 78 78._Leaf_veins / present absent, 79 79._Anther_thecae / tetrasporangiate disporangiate, 80 80._Raphides / absent present, 81 81._Biforines / absent present ; MATRIX [ 10 20 30 40 50 60 70 80] [ . . . . . . . . ] Arum 110111150212000001000012120300022000002000042000001013022100--014142100{01}001000010 Dracunculus 110111150010010001000002122300022000002000042000002013042100--0141421000001000010 Biarum 11011115021201000100001012030002200000200{01}04200000{01}013031101--0041421NNNNN1000010 Helicodiceros 1101111502120N0001000002122300022000002000041000001013042100--0141421000001000010 Eminium 11011115021201N0N100000212230002200000200{01}042000001013032101--0141421NNNNN1000010 Sauromatum 110111150212010001000002122300022000002001042000001013031100--004{12}40201100100001N Typhonium 11011115021001000100000212230002200000200004200000{12}013031101--0042401NNNNN1000010 Theriophonum 11011115021001N0N100001012030002200000200004200000{12}013042100--0041421NNNNN1000010 Pinellia 110111150212010001000002122300022000002100142000002010031001--0142432001001000010 Arisaema 11011115020201000100000212230002200000200003200001{01}013032100--014{12}42100{01}001000010 Alocasia 11011115021201000200101212020001{12}000012000041000121010032000--024{12}432000001000011 Steudnera 110111150003010002001112120200001100012000110000121110022000--0241422001001000010 Remusatia 11011115021201000200111212020012410001200002000012101{23}0{23}2000--0241422NNNNN0000010 Colocasia 11011115001201000200111212020001{12}100012000042000121012022000--024{12}422011NN1000011 Ariopsis 110111150202000001000000120200020100001000120000122013022000--0142402NNNNN1000011 Pistia 110111150013010000000000100300001020002101140010020013032000--024{12}431001111000011 Protarum 1101111500000NN0N2000002122200022000012000042000122112032000--0141NNNNNNNN1000011 Carlephyton 110111150212010001000000120200022000010000110000011113132001--0241422NNNNN1000011 Colletogyne 110111150212010001000000120200022000000000110000000113132001--02414221-0111000011 Arophyton 11011115021201000100000012020000{24}0000{01}200001000001{02}113134001--00424221-0111000011 Typhonodorum 110111150010010001000012120100002010012000041000121113132001--02424221-0{01}{01}1000011 Peltandra 110111150010010001000012120100002010012000041000122112122000--024{12}4221-0001000011 Ambrosina 110111150013010001000000100300022000002101140010020013033000--0041431NNNNN1000010 Arisarum 110111150013010001000010120300012000002001132000010013032000--014{12}421000001000010 Zomicarpa 11011115021201000200000212{24}300022000002000132000002010035000--014{12}422001000000011 Zomicarpella 1101111502N20NN0N2000000100300000000012000132000002010031000--0041401NNNNN0000011 Scaphispatha 1101111502100NN0N2NNNN10120200022100011000010000121010032000--0042122NNNNN1000011 Chlorospatha 110111151210000002110012120200011000012000020000121000001010--0242422NNNNN1000010 Xanthosoma 11011115{01}0100100021100121202000{12}{12}000012000020000121000001010--024{01}421011101000011 Ulearum 1101111502120NN0N2000000100300000000001000142000002010132001--004{12}{34}01NNNNN1000010 Filarum 1101111502020NN0N200000012030002200000100013200000201N1N2001--0041401NNNNN1000010 Syngonium 110111150010010002110012120200003000012000020000121010132001--02414211-0NN1000011 Caladium 110111150211010002110012120200022100012000020000121000022000--0241422011001000011 Hapaline 110111150212010002110012120300022000012000122000121010131001--02414211-0111000011 Jasarum 11011115001001N0N2110010120200002030012000120000122012033000--00524221-0111000010 Pseudodracontium 110111150013010001000011123300022000001000031000001010131001--004{12}422NNNNN1000010 Amorphophallus 1101111500{12}3010001000011123300022000001000031000012000131001--00414221-0111000010 Calla 0101110302000000010000001001000000110000000000-0000-10039-00---25{01}620000000000010 Bucephalandra 1101111500110N0001000010110100001041011000041001002013031000--0001411NNNNN1000010 Aridarum 1101110500010N000100001011010000104101{12}0000{24}{01}0010{01}20120{34}0000--0201021NNNNN1000011 Schismatoglottis 1101111500010{01}00010000101101000010{04}{01}012000041000002{01}10021000--020{12}021011001000011 Phymatarum 1101111500010N0001000010110100001041011000041001002013031000--0201021NNNNN1000010 Piptospatha 1101111500{01}10100010000101101000010410110000{24}{01}000002{01}1{23}0{23}1000--0001011011001000011 Cryptocoryne 110111150011010000000010110100000010102111040001002013039000--0042421000NN1000010 Lagenandra 110111150011010000000010110100010010102111040001002013039000--0041422000001000010 Asterostigma 110111150010010001000010111300022000002000010000122100038001--105{01}022NNNNN1000010 Taccarum 11011115002001000100001111330002200001200001000012{12}100008001--1002022NNNNN1000010 Spathicarpa 110111150011010001000010100300012000000000210000122113038001--1001102001001000010 Synandrospadix 110111150212010001000010110300022000001000120000120103038001--1001022000001000010 Croatiella 1101111500110NNNNNNNNN121203000220000010001100001{12}0123N38001--0NNNNNNNNNNN100001N Gorgonidium 1101111500100N00010000101113000220000110000100001{01}{12}103038001--1001NNNNNNNN1000010 Spathantheum 11011115002101000000001211030002200000100011000012{02}122038001--1001022000001000010 Incarum 1101111500100NNNNNNNNN10111300022000001000110000110123038001--0NNNNNNNNNNN100001N Mangonia 1101111500100NN0N000001010030002200000200003000011{12}100008001--000{01}021000001000010 Dieffenbachia 1101111500200100010000101001000010{01}0002000120000121100138001--02111311-011{01}000011 Gearum 1101111500200NN0N0000012121300012000002000020000122103138001--00010211-0111000011 Bognera 1101111500201NN0N1000010110100000000001000120000120010N38001--0NNNNNNNNNNN100001N Zantedeschia 1101111500110100010000101101000{12}20{01}0002000010000102{01}00007000--0212122001001000011 Furtadoa 1101111500010N1111000010110100001040001000020000100110031000--100{01}001NNNNN1000010 Homalomena 110111150001011011000010110100001000001000020000101100001000--0201001NNNNN1000011 Philodendron 1101111500111111110000101101000010{01}010200002000010{12}000009000--0201001011001000011 Culcasia 110111050210111110000010110300003000021000010000101000131001--01000201-0001000011 Cercestis 110111150010111111000010110300003000002000010000101010131001--01100201-0111000010 Anchomanes 110111150021110001000011120311012000001000010000101010131001--00012311-0111000011 Pseudohydrosme 11011115003101N0N0000011120311022000002000010000102000131001--0021NNN1-0111000011 Nephthytis 11011115001011000100000112430{01}012000000000010000102010131001--01012011-0111000011 Aglaodorum 110111150021010001000010100100000010021000010000102100121001--0021221NNNNN1000011 Aglaonema 110111150021010001000010100100001000021000010000102010131000--10212211-0111000011 Montrichardia 11011115022100100100001111030{01}002010022000010000101010130001--020{01}6221-0111000010 Anubias 110111150201101001000010110100000010001000010000120000000000--0200{03}01011001000011 Callopsis 110111150212010001000010100300002000000000110000002010039001--005{01}000000001000010 Gonatopus 100111110020110000000010115300012000002000010000012000138001011051{03}011-0110000010 Zamioculcas 100111110020110000000010111300012000002000010000000000138001011000001NNNNN0000010 Stylochaeton 100111150020010000000010100300002000002001010000000000002000111201011NNNNN0000011 Dracontioides 0001111000001000000000011043100020100020000001-0001-00001-0101-050631000000000010 Anaphyllopsis 0001111000100N00000000111233100220000010000001-0001-10031-0101-050NNNNNNNN0000010 Dracontium 0001111000101000000000111233100220000010000001-0001-00001-0101-00{01}031NNNNN0000010 Urospatha 0001111000101000000000111143000120100010000001-0001-00001-0001-000030000000000010 Podolasia 0001111000001N00000000011043010010100000000001-0000-10031-0101-350630NNNNN0000010 Pycnospatha 0101111000100NN0N00000111133100220000010000001-0000-10031-01---300631NNNNN0000010 Cyrtosperma 0001111000101000000000111143010120100010000001-0000-11021-0001-300030000000000010 Anaphyllum 0001111000101N00000000111133000000{01}00000000000-0001-10121-0101-351030NNNNN0000010 Lasimorpha 0001111000001NNNN00000111143010120100010000001-0011-11031-0001-300030NNNNN0000010 Lasia 0001111000101N00000000121223010010100210000001-0000-10011-0101-300030NNNNN0000010 Amydrium 010111N201101{01}00000000101103100030000210000000-0000-10036-01---101{03}00NNNNN0000010 Epipremnum 0111110201101100000000101101100030000210000000-0000-10026-00---050{03}00NNNNN0000010 Monstera 0111110201101100000000101101100030000210000000-0000-00136-00---051{03}111-0000000010 Scindapsus 0111110201101{01}00000000101101000030000210000000-0000-10036-01---00{01}{03}00000000000010 Anadendrum 0001110502001N00000000101103000030000210000000-0000-10136-0110-000{35}001-0100000010 Rhaphidophora 0111110100101N00000000101101100030000210000000-0000-00006-00---05{01}{03}00NNNNN0000010 Holochlamys 0021110900131000000000101101000000000000000000-0000-10036-0110-101001NNNNN0000010 Spathiphyllum 0021110900131000000000101101000000000000000000-0000-00006-0000-000011001000000010 Alloschemone 0111110200101NNNN00000101101100030000210000000-0000-10N32-01---101610NNNNN0000010 Rhodospatha 0111110200101100000000101101000030000210000000-0000-00002-00---000610NNNNN0000010 Heteropsis 0101130200101100000000101203000030000210000000-0001-00132-01---0506001-0000000010 Stenospermation 0111110201101100000000101101000030000210000000-0000-00032-00---05{01}011000000000010 Pothoidium 0001130000101N00000000001000000030000000000000-0000-10130-0100-050600NNNNN0000010 Pedicellarum 0001130000001NNNN00000001000000030000000000000-0000-00130-0110-050600NNNNN0000010 Pothos 0001130000001N00000000001000000030000000000000-0000-00130-0100-1506001-0000000010 Anthurium 0001111402001000000000001000000030000000000000-0000-00006-0101-10000000{01}NN0000010 Wolffia 0102---70202-N-N-0------2-----03--2-0----------0000-13035-01---25N6--NNNNN01-1110 Wolffiella 0102---70202-N-N-0------2-----03--2-0----------0000-13035-01---25N6--NNNNN01-1110 Landoltia 0100---70202-N0N00----0010040-03-0200----------0000-10035-01---N5N6--0011100100NN Lemna 0100---70202-N0N00----0010040-03-0200----------0000-10035-0{01}---25N6--001110010010 Spirodela 0100---70202-N0N00----0010040-03-0200----------0000-10035-01---25N6--001110010010 Symplocarpus 0001101000100000000000101000000120100010000000-0000-13116-0100-200{03}201-0000000010 Lysichiton 0001001000100100000000101000000120100010000000-0000-03102-0100-2303201-0000000010 Orontium 000000N000200000010000001000000120100000000000-0000-10131-0100-2506201-0000000010 Gymnostachys 0000020000100000000000000000000120000000000000-0000-13010-0100-050600000000000010 Tofieldia 00000-0302{01}00N00000000000001000020100000000000-0000-0{01}006-0000-NNNNNNNNNNN000000- Acorus 0000000000000000000000000000000000101000000000-0000-03010-0000-25062000100000000- ; END; BEGIN mrbayes; outgroup Acorus; lset coding=variable; mcmcp ngen=10000000 printfreq=100 samplefreq=100 nchains=4 savebrlens=yes filename=Cusimanoetal_Morpho_10mio; mcmc; END; BEGIN LABELS; CharGroupLabel group_2 COLOR=(RGB 0.73307 0.73307 0.73307 ); END; BEGIN SETS; CHARSET Good_characters = 1 3 8 12 16-18 46 49-50 57-58 64-65 67-68 75; TAXSET Areae = 1-8; TAXSET Colocasia_clade = 12-15; TAXSET Amorphophallus_clade = 25-37; END; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ; END; BEGIN TREES; TREE * MLtree = [&R] 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