Biarum pyrami (Araceae)
Tuber globose-discoid 2–4 × 2–2.5 cm, mid-brown. Leaves hysteranthous (var. pyrami ) or synanthous (var. serotinum ), long-petiolate, encased basally by numerous (2 – )7–13 × 0.5–2 cm elliptic-lanceolate sub-fleshy, later papery, cataphylls, these pale greenish, occasionally with faint purple spots externally at first, later pale straw-yellow; petiole 8–16 cm × 2.5–4.5 mm, margins slightly winged, adaxial surface channeled distally, expanded proximally into broad membranous wing, mid-green; leaf lamina oblong-elliptic to ovateelliptic, 6–18 × 3–8 cm, apex subacute to slightly rounded, base briefly decurrent to rounded, occasionally sub-truncate, c. 9–10 primary lateral veins per side, margins fl at, occasionally crispulate, leaf lamina deep green, with black-purple or green bullae (var. pyrami ) or smooth (var. serotinum ). Inflorescence appearing in late autumn (var. pyrami ) or early winter (var. serotinum ), smelling strongly foetid of cattle dung and carrion; peduncle 4–7 cm × 3–4 mm, off white. Spathe 10–25 cm long; spathe limb lanceolate, (7 – )15–18.5 × 2–3.5 cm, long-acuminate, erect at first but soon refl exing and coiling, exterior mid-green stained and spotted purple-brown, especially towards the margins (var. pyrami ) or unspotted (var. serotinum ), interior deep purple brown, sometimes with paler speckling along the middle and towards the apex; spathe tube globose, ventricose, 3–4 × 2.5–3.5 cm, margins connate for a ¼ of their length, exterior whitish below ground, purple-brown above, interior dark purple. Spadix equalling to slightly exceeding the spathe limb, 9–26 cm long; spadix appendix cylindricfusiform 5.7–22 cm × (3 – ) 5–9 mm, deep purple-brown. Staminate flowers in a zone 10–15 × 5–6 mm, anthers purple. Interstice 14–16 × 2–3.5 mm, deep purple, rarely dull creamy yellow. Pistillodes in a zone 5–10 mm long at the base of the interstice, sparse; bases slightly swollen, purple; bristles filiform, flattened distally, 5–7 mm long, cream, sometimes flushed deep purple. Pistillate flowers in a hemispherical cluster c. 10 × 3–5 mm high, ovaries bottle-shaped, 1.5–2 × 1–1.25 mm, deep purple, paler below, style c. 1.5 × 0.33 mm, purple, stigma sub-capitate, c. 0.50 mm in diam., white. Infructescence globose, 2–3 cm in diam., consisting of c. 50 berries; berries oblong-pyriform, 7–10 × 5–6 mm, silvery-lilac, stained purple proximally, seed sub-turbinate, 5–6 × 4–5 mm wide, testa reticulate, dark brown.
This large, showy species is perhaps the most readily distinguishable of the autumn flowering Biarum species due to the globose, strongly inflated, spathe tube, large spathe and greatly attenuated spadix appendix. The bullate leaves of the typical variety are also unusual in the genus and provide a ready means of identifying B. pyrami in Turkey, where no other species shares this character.