Albertarum

General description: 

FOSSIL

GENERIC DESCRIPTION: Infructescence stipitate, subglobose. Flowers bisexual, perigoniate; tepals six, fornicate, imbricate. Gynoecium with a trilocular ovary, one ovule per locule, placenta apical-parietal or possibly mid-axial; ovules
anatropous; stylar region long attenuate. Seed ellipsoid; testa thick, prominently ribbed; raphe present.

Diagnostic description: 

DIAGNOSIS: Albertarum pueri a genere extanti Symplocarpo, flore 6 tepalis instructo, gynoecii ovario 3-loculare, loculis
uniovulatis, ovulo anatropo, semine ellipsoideo, testa crassa et costata, raphe distincta differt.

Taxonomy
Taxonomic Notes: 

Affinity
The fossil flowers are bisexual and perigoniate. Bisexual flowers occur in subfamilies Gymnostachydoideae, Orontioideae, Pothoideae, Monsteroideae, Lasioideae, and Calloideae. In Gymnostachydoideae, Orontioideae, and Pothoideae, all genera are perigoniate. In Monsteroideae, only three genera (Spathiphyllum Schott, Holochlamys Engler, and Anadendrum Schott) have a perigone (Engler and Krause 1908). In Lasioideae, all genera except one (Pycnospatha Thorel ex Gagnepain) have a perigone. The single genus of Calloideae, Calla L., lacks a perigone (Krause 1908). The fossil is referred to subfamily Orontioideae. It is excluded from subfamily Calloideae by its perigone and from subfamilies Gymnostachydoideae, Pothoideae, Monsteroideae, and Lasioideae by the morphology of the stylar region. In those subfamilies, only some species of Spathiphyllum have a similar stylar region, but they have a cylindric rather than globose infructescence, and their seeds are not ribbed like those of the fossil (Seubert 1993). There are three extant genera in subfamily Orontioideae: Symplocarpus, Lysichiton Schott, and Orontium L. (Mayo et al. 1997). The fossil shares a few characters with each of them, but there are significant differences (Table 1); therefore, the new genus Albertarum is erected for the fossil. Albertarum is most similar to Symplocarpus (Table 1). The morphology of the stylar region is quite similar, and both have a globose to subglobose infructescence (Figs. 8, 9). Albertarum is distinguished by its trilocular ovary, however, and its ellipsoid, ribbed seeds. It also differs in the number of tepals, and the anatropous ovules.

Taxonomic name: 
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith