Bucephalandra muluensis
Small obligate rheophytic herbs to 10 cm tall. Stem initially erect, later creeping with active portion sub-erect, c. 5 mm in diam. Leaves several together; petiole 2 - 3 cm long × c. 1.5 mm in diam., adaxially canaliculate, bright green, sheathing at extreme base, wings extended into a very narrowly triangular ligular portion to 1 cm long; blade ovate to broadly elliptic, 2 - 4 cm long × 1.5 - 2 cm wide, thinly coriaceous, weakly glossy bright green, slightly paler abaxially, base rounded to rounded-cuneate, apex acute and apiculate for c. 2 mm, margin flat; midrib abaxially and adaxially sharply prominent; primary lateral veins 3 or 4 on each side, diverging at c. 40° and running to a weak marginal vein; interprimary veins very weak; all other venation obscure. Inflorescence(s) solitary to up to 3 together interspersed with prophylls; peduncle exceeding petioles, c. 4 cm long, sub-terete, pale green. Spathe broadly ovate, not constricted, c. 3 cm long; lower spathe funnel-form, yellowish green, persistent; limb inflating and gaping distally at pistillate anthesis, then deliquescent-caducous during staminate anthesis, glistening white, apiculate for c. 2 mm, apiculum green. Spadix 1.3 - 1.5 cm long; pistillate zone c. 3 mm long × c. 1.5 mm in diam., with c. 3 spirals of pistils; pistils rhombic-globose, c. 0.4 mm in diam., bright green; stigma sessile, umbonate, c. 2/3 diameter of ovary, papillate at anthesis; pistillodes 2 - 4 at base of pistillate zone, bottle-shaped, c. 0.35 mm long., white, tip green; interstice with c. 2 rows of scale-like staminodes, these 1 - 2 mm long × 0.8 - 1.5 mm wide, spathulate, initially erect, then spreading, initially matte white, later (post-anthesis) becoming green and persisting to cover developing fruit; staminate zone c. 3 mm long × c. 2 mm in diam., consisting of c. 4 rows of flowers; staminate flowers medium yellow; stamen comparatively large, c. 1 mm across; filament short, slender; connective slender, prismatic; thecae inserted ventrally, ellipsoid, c. 1 mm long × c. 0.4 mm wide, smooth; thecae horns sub-equalling associated theca, setaceous, spreading; appendix ellipsoid, basally somewhat abruptly transitioning to staminate flower zone, 7 - 9 mm long × 3 - 4 mm in diam., medium yellow; appendix staminodes individually well-demarcated, obpyramidal, 0.5 - 1 mm in diam., upper surface brain-like with a central impression. Fruiting spathe funnel-form, c. 1 cm in diam., with shield shaped staminodes persistent, turning green, spent distal part of spadix falling; berry depressed globose, 1 - 1.8 mm long × 1 - 1.5 mm in diam., with numerous seeds, berries forming a hemispherical cluster protected by persistent staminodes, later pushed off and falling to reveal mature fruits.
Known only on the Mulu National Park limestones, where it is scattered and not common.
Bucephalandra muluensis is rheophytic on exposed to semi-shady karst limestone rocks along streams under perhumid to moist lowland forest, below 100 m above sea level.