Schismatoglottis petradoxa
Pendent rheophytic herb to ca. 50 cm. Stem much-condensed, creeping-erect in older plants, with internodes to 1 cm long, 1–2 cm diam., densely rooting along its length. Leaves several together alternating soon-marcescent, somewhat brittle tapering lanceolate weakly scabrid cataphylls to 10 cm long; petiole shorter than blade, arching to almost pendent, 8–25 cm long, subterete, proximal third dorsally very slightly flattened, dull green, minutely scabrid, sheathing only at very base, with the petiolar sheath reduced to an obscure ridge; blade broadly lanceolate to elliptic, 12–30 cm long × 3–11 cm wide, thick, brittle, adaxially matte medium green, much paler and matte abaxially, base cuneate to narrowly rounded, apex acuminate and apiculate for ca. 1 cm; midrib adaxially more or less flush with blade and contrasting cream, especially visible on newer leaves, abaxially prominent; primary lateral veins ca. 25 on each side, up to 40 on largest leaf blades, diverging at 45–60°, abaxially conspicuously raised, adaxially slightly impressed, although more deeply impressed in leaves of seedlings; secondary venation adaxially visible as a tessellate reticulum in seedlings but obscure in leaves of mature plants, abaxially forming a highly conspicuous dark green tessellate reticulum; tertiary venation invisible. Inflorescence pendulous, ca. 3 in a simple synflorescence subtended by lanceolate fleshy cataphylls, with a moderate sweet esteric odour at pistillate anthesis; peduncle cylindric, subtended by a conspicuous prophyll or cataphyll, up to 20 cm long × 5 mm wide, bright pale green. Spathe with a moderate constriction between the lower part and the limb, limb inflating and gaping at pistillate anthesis, opening further at staminate anthesis, 8–9 cm long; lower spathe narrowly ovoid and asymmetric, dorsally shallowly flattened-convex corresponding to the adnation of the pistillate flower zone, pale green with very fine darker longitudinal veins, dorsally ca. 1.5 cm long, ventrally ca. 2 cm long, persistent; spathe limb exterior white with very fine darker longitudinal lines at pistillate anthesis, interior dirty whitish-green shading dorsally to glossy green broadly lanceolate 5.5–7.5 cm long, bluntly rostrate for ca. 1 cm, upper half opening at pistillate anthesis and initially via a narrow terminal slit, then wide-gaping (ca. 2.5 cm wide), and weakly fornicate, limb margins reflexing during staminate anthesis, then whole limb degrading-caducous with the rim remaining above the lower spathe insertion reflexing somewhat. Spadix 5–6 cm long, subcylindric; pistillate zone 1.5 (dorsal side) to 2 cm long (ventral side), narrowly conic, obliquely inserted, distally ca. 6 mm diam., very pale green; pistils small, crowded, ca. 1 mm diam., barrel-shaped, pale creamy green; stigma sessile, discoid, slightly narrower than top of pistil, ca. 1.5 mm tall × 0.8 mm wide, papillose; interpistillar pistillodes forming a row at junction with peduncle, ca. 1.2 mm long, slimmer than pistils, slender-cylindric with a pale orange-brown tip; sterile interstice with about 3 rows of staminodes; interstice staminodes weakly columnar-polygonal ca. 1 mm across, dull-white, initially equalling the height of pistils, later (late pistillate anthesis) staminodes lengthening to form a ring ca 1/3 wider than fertile zones; staminate zone cylindric, ca. 1.5 cm long × 0.4–0.5 cm diam., cream; stamens irregularly densely crowded, individual flowers difficult to distinguish, rectangular-dumbbell shaped from above, truncate with thick connective slightly elevated above thecae, thecae opening by a single pore; appendix narrowly fusiform, blunt, proximally slightly wider than top of staminate zone, 2.5–3 cm long, widest part ca. 8 mm diam., distally tapering and narrowly obtuse, white; appendix staminodes very dense and individually barely differentiated, rectangular-dumbbell shaped from above, much resembling stamens in shape and size, but more regularly arranged. Fruiting spadix not seen.
Schismatoglottis petradoxa is defined by the combination of condensed pleionanthic shoots composed of many single leaved modules, each subtended by a soon-degrading prophyll, leaf blades abaxially with conspicuous tessellate secondary venation, and a synflorescence of pendent inflorescences, each on a long peduncle. In the shoot organization S. petradoxa is reminiscent of species of the Schismatoglottis Tecturata Group, but otherwise differs by the above mentioned characteristics.
Widespread but localized in central North Borneo.
Schismatoglottis petradoxa occurs as a rheophyte on shaded vertical permanently wet shale waterfalls and rocks in forest streams between 30 – 190 m a.s.l.